<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE article PUBLIC "-//NLM//DTD JATS (Z39.96) Journal Publishing DTD v1.3 20210610//EN" "JATS-journalpublishing1-3.dtd">
<article article-type="research-article" dtd-version="1.3" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xml:lang="en"><front><journal-meta><journal-id journal-id-type="publisher-id">nsc</journal-id><journal-title-group><journal-title xml:lang="en">Archaeology, Ethnology &amp; Anthropology of Eurasia</journal-title><trans-title-group xml:lang="ru"><trans-title>Археология, этнография и антропология Евразии</trans-title></trans-title-group></journal-title-group><issn pub-type="ppub">1563-0110</issn><publisher><publisher-name>IAET SB RAS</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="doi">10.17746/1563-0110.2026.54.2.027-036</article-id><article-id custom-type="elpub" pub-id-type="custom">nsc-2467</article-id><article-categories><subj-group subj-group-type="heading"><subject>Research Article</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="en"><subject>PALEOENVIRONMENT. THE STONE AGE</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="ru"><subject>ПАЛЕОЭКОЛОГИЯ. КАМЕННЫЙ ВЕК</subject></subj-group></article-categories><title-group><article-title>Seasonal Cycles and Behavioral Strategies of the Paleolithic Dwellers in Denisova Cave</article-title><trans-title-group xml:lang="ru"><trans-title></trans-title></trans-title-group></title-group><contrib-group><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="western" xml:lang="en"><surname>Shunkov</surname><given-names>M. V.</given-names></name></name-alternatives><bio xml:lang="en"><p>630090; Pr. Akademika Lavrentieva 17; Novosibirsk</p></bio><email xlink:type="simple">shunkov77@gmail.com</email><xref ref-type="aff" rid="aff-1"/></contrib><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="western" xml:lang="en"><surname>Agadjanian</surname><given-names>A. K.</given-names></name></name-alternatives><bio xml:lang="en"><p>117868; Profsoyuznaya 123; Moscow</p></bio><email xlink:type="simple">aagadj@paleo.ru</email><xref ref-type="aff" rid="aff-2"/></contrib></contrib-group><aff xml:lang="en" id="aff-1"><institution>Institute of Archaeology and Ethnography, Siberian Branch, Russian Academy of Sciences</institution><country>Russian Federation</country></aff><aff xml:lang="en" id="aff-2"><institution>Borissiak Paleontological Institute, Russian Academy of Sciences</institution><country>Russian Federation</country></aff><pub-date pub-type="collection"><year>2026</year></pub-date><pub-date pub-type="epub"><day>26</day><month>07</month><year>2026</year></pub-date><volume>54</volume><issue>2</issue><fpage>27</fpage><lpage>36</lpage><permissions><copyright-statement>Copyright &amp;#x00A9; Shunkov M.V., Agadjanian A.K., 2026</copyright-statement><copyright-year>2026</copyright-year><copyright-holder xml:lang="ru">Shunkov M.V., Agadjanian A.K.</copyright-holder><copyright-holder xml:lang="en">Shunkov M.V., Agadjanian A.K.</copyright-holder><license xml:lang="ru" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>Данная работа распространяется под лицензией Creative Commons Attribution 4.0.</license-p></license><license xml:lang="en" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>This work is licensed under a Creative Commons Attribution 4.0 License.</license-p></license></permissions><self-uri xlink:href="https://journal.archaeology.nsc.ru/jour/article/view/2467">https://journal.archaeology.nsc.ru/jour/article/view/2467</self-uri><abstract><p>   A model of seasonal cycles in the subsistence strategy of Middle and Upper Pleistocene humans in and around Denisova Cave, the Anuy River valley, northwestern Altai, is proposed. On the basis of the year-round monitoring of temperature changes in the cave and around it, certain regularities of circadian and seasonal fluctuations are evaluated. Findings suggest that the climatic situation in winter was more comfortable in the cave than in the Anuy valley, whereas in summer the opposite was true. Faunal data in the cave indicate competition between Paleolithic humans and large carnivores. The main rivals at the early stage were bears, which used the karst cave for their dens in winter. As human activities intensified, hyenas, who needed the cave for breeding in spring and summer, became the principal competitors (judging by the abundance of bones and coprolites), and their relationship with humans varied (alternating or seasonal) over many years. Findings suggest that Paleolithic humans lived in the cave mostly in autumn and winter, shifting to nearby open-air camps in spring and summer. The use of camps on stepped slopes in the extensions of the valley provided much better conditions for hunting.</p></abstract><kwd-group xml:lang="en"><kwd>Altai</kwd><kwd>Denisova Cave</kwd><kwd>Anuy valley</kwd><kwd>Pleistocene</kwd><kwd>temperature regime</kwd><kwd>human paleoecology</kwd></kwd-group><funding-group><funding-statement xml:lang="en">The study was supported by the Russian Science Foundation, Grant No. 24-18-00069, https://rscf.ru/project/24-18-00069/</funding-statement></funding-group></article-meta></front><back><ref-list><title>References</title><ref id="cit1"><label>1</label><citation-alternatives><mixed-citation xml:lang="ru">Agadjanian A.K., Serdyuk V.N. 2005 The history of mammalian communities and paleogeography of Altai Mountains in the Paleolithic. Paleontological Journal, vol. 39, suppl. 6: 645–821.</mixed-citation><mixed-citation xml:lang="en">Agadjanian A.K., Serdyuk V.N. 2005 The history of mammalian communities and paleogeography of Altai Mountains in the Paleolithic. Paleontological Journal, vol. 39, suppl. 6: 645–821.</mixed-citation></citation-alternatives></ref><ref id="cit2"><label>2</label><citation-alternatives><mixed-citation xml:lang="ru">Derevianko A.P. 2022 Three Global Human Migrations in Eurasia, vol. VI. Pt. 1: Denisovan Human: Origins, Material and Spiritual Culture. Novosibirsk: Izd. IAET SO RAN, 900 p.</mixed-citation><mixed-citation xml:lang="en">Derevianko A.P. 2022 Three Global Human Migrations in Eurasia, vol. VI. Pt. 1: Denisovan Human: Origins, Material and Spiritual Culture. Novosibirsk: Izd. IAET SO RAN, 900 p.</mixed-citation></citation-alternatives></ref><ref id="cit3"><label>3</label><citation-alternatives><mixed-citation xml:lang="ru">Derevianko A.P., Shunkov M.V. 2002 Middle Paleolithic industries with foliate bifaces in Gorny Altai. Archaeology, Ethnology and Anthropology of Eurasia, No. 1: 16–41. (In Russian and English).</mixed-citation><mixed-citation xml:lang="en">Derevianko A.P., Shunkov M.V. 2002 Middle Paleolithic industries with foliate bifaces in Gorny Altai. Archaeology, Ethnology and Anthropology of Eurasia, No. 1: 16–41. (In Russian and English).</mixed-citation></citation-alternatives></ref><ref id="cit4"><label>4</label><citation-alternatives><mixed-citation xml:lang="ru">Derevianko A.P., Shunkov M.V., Agadjanian A.K., Baryshnikov G.F., Malaeva E.M., Ulyanov V.A., Kulik N.A., Postnov A.V., Anoikin A.A. 2003 Prirodnaya sreda i chelovek v paleolite Gornogo Altaya. Novosibirsk: Izd. IAET SO RAN, 448 p.</mixed-citation><mixed-citation xml:lang="en">Derevianko A.P., Shunkov M.V., Agadjanian A.K., Baryshnikov G.F., Malaeva E.M., Ulyanov V.A., Kulik N.A., Postnov A.V., Anoikin A.A. 2003 Prirodnaya sreda i chelovek v paleolite Gornogo Altaya. Novosibirsk: Izd. IAET SO RAN, 448 p.</mixed-citation></citation-alternatives></ref><ref id="cit5"><label>5</label><citation-alternatives><mixed-citation xml:lang="ru">Jacobs S., Hudjashov G., Saag L., Kusuma P., Darusallam C.C., Lawson D.J., Mondal M., Pagani L., Ricaut F.-X., Stoneking M., Metspalu M., Sudoyo H., Stephen L.J., Cox M.P. 2019 Multiple deeply divergent Denisovan ancestries in Papuans. Cell, vol. 177 (4): 1010–1021.</mixed-citation><mixed-citation xml:lang="en">Jacobs S., Hudjashov G., Saag L., Kusuma P., Darusallam C.C., Lawson D.J., Mondal M., Pagani L., Ricaut F.-X., Stoneking M., Metspalu M., Sudoyo H., Stephen L.J., Cox M.P. 2019 Multiple deeply divergent Denisovan ancestries in Papuans. Cell, vol. 177 (4): 1010–1021.</mixed-citation></citation-alternatives></ref><ref id="cit6"><label>6</label><citation-alternatives><mixed-citation xml:lang="ru">Krause J., Fu Q., Good J.M., Viola B., Shunkov M.V., Derevianko A.P., Pääbo S. 2010 The complete mitochondrial DNA genome of an unknown hominin from Southern Siberia. Nature, vol. 464: 894–897.</mixed-citation><mixed-citation xml:lang="en">Krause J., Fu Q., Good J.M., Viola B., Shunkov M.V., Derevianko A.P., Pääbo S. 2010 The complete mitochondrial DNA genome of an unknown hominin from Southern Siberia. Nature, vol. 464: 894–897.</mixed-citation></citation-alternatives></ref><ref id="cit7"><label>7</label><citation-alternatives><mixed-citation xml:lang="ru">Kulik N.A., Shunkov M.V. 2000 Predvaritelniye rezultaty petrografi cheskogo izucheniya paleoliticheskikh izdeliy stoyanki Anuy-3. In Problemy arkheologii, etnografii, antropologii Sibiri i sopredelnykh territoriy, vol. VI. Novosibirsk: Izd. IAET SO RAN, pp. 156–160.</mixed-citation><mixed-citation xml:lang="en">Kulik N.A., Shunkov M.V. 2000 Predvaritelniye rezultaty petrografi cheskogo izucheniya paleoliticheskikh izdeliy stoyanki Anuy-3. In Problemy arkheologii, etnografii, antropologii Sibiri i sopredelnykh territoriy, vol. VI. Novosibirsk: Izd. IAET SO RAN, pp. 156–160.</mixed-citation></citation-alternatives></ref><ref id="cit8"><label>8</label><citation-alternatives><mixed-citation xml:lang="ru">Morley M.V., Goldberg P., Uliyanov V.A., Kozlikin M.B., Shunkov M.V., Derevianko A.P., Jacobs Z., Roberts R.G. 2019 Hominin and animal activities in the microstratigraphic record from Denisova Cave (Altai Mountains, Russia). Scientific Reports, vol. 9 (13785): 1–12.</mixed-citation><mixed-citation xml:lang="en">Morley M.V., Goldberg P., Uliyanov V.A., Kozlikin M.B., Shunkov M.V., Derevianko A.P., Jacobs Z., Roberts R.G. 2019 Hominin and animal activities in the microstratigraphic record from Denisova Cave (Altai Mountains, Russia). Scientific Reports, vol. 9 (13785): 1–12.</mixed-citation></citation-alternatives></ref><ref id="cit9"><label>9</label><citation-alternatives><mixed-citation xml:lang="ru">Postnov A.V. 1999 Metod levalluazskogo rasshchepleniya, rekonstruirovanniy na osnove remontazha artefaktov stoyanki Ust-Karakol-1 (Gorniy Altai). Gumanitarniye nauki v Sibiri, No. 3: 16–20.</mixed-citation><mixed-citation xml:lang="en">Postnov A.V. 1999 Metod levalluazskogo rasshchepleniya, rekonstruirovanniy na osnove remontazha artefaktov stoyanki Ust-Karakol-1 (Gorniy Altai). Gumanitarniye nauki v Sibiri, No. 3: 16–20.</mixed-citation></citation-alternatives></ref><ref id="cit10"><label>10</label><citation-alternatives><mixed-citation xml:lang="ru">Prüfer K., Racimo F., Patterson N., Jay F., Sankararaman S., Sawyer S., Heinze A., Renaud G., Sudmant P.H., Filippo C., de, Heng Li, Mallick S., Dannemann M., Qiaomei Fu, Kircher M., Kuhlwilm M., Lachmann M., Meyer M., Ongyerth M., Siebauer M., Theunert C., Tandon A., Moorjani P., Pickrell J., Mullikin J.C., Vohr S.H., Green R.E., Hellmann I., Johnson P.L.F., Blanche H., Cann H., Kitzman J.O., Shendure J., Eichler E.E., Lein E.S., Bakken T.E., Golovanova L.V., Doronichev V.B., Shunkov M.V., Derevianko A.P., Viola B., Slatkin M., Reich D., Kelso J., Pääbo S. 2014 The complete genome sequence of a Neanderthal from the Altai mountains. Nature, vol. 505 (7481): 43–49.</mixed-citation><mixed-citation xml:lang="en">Prüfer K., Racimo F., Patterson N., Jay F., Sankararaman S., Sawyer S., Heinze A., Renaud G., Sudmant P.H., Filippo C., de, Heng Li, Mallick S., Dannemann M., Qiaomei Fu, Kircher M., Kuhlwilm M., Lachmann M., Meyer M., Ongyerth M., Siebauer M., Theunert C., Tandon A., Moorjani P., Pickrell J., Mullikin J.C., Vohr S.H., Green R.E., Hellmann I., Johnson P.L.F., Blanche H., Cann H., Kitzman J.O., Shendure J., Eichler E.E., Lein E.S., Bakken T.E., Golovanova L.V., Doronichev V.B., Shunkov M.V., Derevianko A.P., Viola B., Slatkin M., Reich D., Kelso J., Pääbo S. 2014 The complete genome sequence of a Neanderthal from the Altai mountains. Nature, vol. 505 (7481): 43–49.</mixed-citation></citation-alternatives></ref><ref id="cit11"><label>11</label><citation-alternatives><mixed-citation xml:lang="ru">Rohland N., Pollack J.L., Nagel D., Beauval C., Airvaux J., Paabo S., Hofreiter M. 2005 The population history of extant and extinct hyenas. Molecular Biology and Evolution, vol. 22: 2435–2443.</mixed-citation><mixed-citation xml:lang="en">Rohland N., Pollack J.L., Nagel D., Beauval C., Airvaux J., Paabo S., Hofreiter M. 2005 The population history of extant and extinct hyenas. Molecular Biology and Evolution, vol. 22: 2435–2443.</mixed-citation></citation-alternatives></ref><ref id="cit12"><label>12</label><citation-alternatives><mixed-citation xml:lang="ru">Slon V., Mafessoni F., Vernot B., Filippo C., de, Grote S., Viola B., Hajdinjak M., Peyrégne S., Nagel S., Brown S., Douka K., Higham T., Kozlikin M.B., Shunkov M.V., Derevianko A.P., Kelso J., Meyer M., Prüfer K., Pääbo S. 2018 The genome of the offspring of a Neandertal mother and a Denisovan father. Nature, vol. 561 (7721): 113–116.</mixed-citation><mixed-citation xml:lang="en">Slon V., Mafessoni F., Vernot B., Filippo C., de, Grote S., Viola B., Hajdinjak M., Peyrégne S., Nagel S., Brown S., Douka K., Higham T., Kozlikin M.B., Shunkov M.V., Derevianko A.P., Kelso J., Meyer M., Prüfer K., Pääbo S. 2018 The genome of the offspring of a Neandertal mother and a Denisovan father. Nature, vol. 561 (7721): 113–116.</mixed-citation></citation-alternatives></ref></ref-list><fn-group><fn fn-type="conflict"><p>The authors declare that there are no conflicts of interest present.</p></fn></fn-group></back></article>
